Similar to Assassin's Creed: Altaïr's Chronicles, the environment is a key feature of gameplay. To keep his temperature warm, Altaïr must approach the torches spread throughout several locations. When the temperature bar hits its bottom, Altaïr will die of freezing. Special to the gameplay in the cold, snowy Masyaf is the temperature bar, a vertical bar on the right side of the screen. Altaïr's health is indicated by a health bar at the top left corner of the screen, consisting of three sections at the beginning of the game but gradually expanding as the player discovers hidden memory blocks. To traverse them, Altaïr has various abilities, including running, dashing, crouching, rolling, jumping, swinging on poles or chains, blending with scholars, and attacking with his sword, grappling hook, bombs, and crossbow. The game is composed of 13 levels (8 in the low-end version) set in Acre, Masyaf, and Jerusalem. With Sinan dead, Altaïr obtains and hides the treasure he had previously looted from Robert de Sablé: the elusive Piece of Eden capable of altering the world's fate. With this revelation of Sinan's betrayal of the Creed, Altaïr is forced to confront and kill his former master. With his reputation restored and his missions completed, Altaïr returns to Sinan, only to discover that Sinan is in fact secretly a Templar who had manipulated Altaïr to eliminate Sinan's former allies to consolidate more power for himself. He first kills Sibrand, William de Montferrat, and Garnier de Naplouse in Acre, then strikes Vizier Abull Aswad and Tamir Bin Musa near Masyaf, and finishes with Maj Aldin, Tallal, and Robert de Sablé in Jerusalem. Altaïr is ordered to kill seven high-ranking Templars scattered throughout the three cities of Acre, Masyaf, and Jerusalem. After he failed to assassinate Robert de Sablé and led the Templars back to Masyaf, he was demoted, though Altaïr's master and the Assassins' leader Sinan offered him a task to regain his former reputation. Before the Third Crusade, he is the most skilled Assassin in the entire Order. Altaïr is a member of the Assassin Order, a faction in opposition to the Knights Templar.

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